위키백과

등골나물족

등골나물족(----族, 학명: Eupatorieae 에우파토리에아이[*])은 국화아과의 족이다.[1] 약 2,000여 종 이상을 포함하고 있다.

등골나물족
카나비눔등골나물(Eupatorium cannabinum)
카나비눔등골나물(Eupatorium cannabinum)
생물 분류ℹ️
계: 식물계
(미분류): 속씨식물군
(미분류): 진정쌍떡잎식물군
(미분류): 국화군
목: 국화목
과: 국화과
아과: 국화아과
족: 등골나물족
Eupatorieae
Cass., 1819
모식종
등골나물속(Eupatorium L.)
[출처 필요]
속

하위 분류 편집

깁티스아족(Gyptidinae R.M.King & H.Rob.)[2]
  • Agrianthus Mart. ex DC.
  • Arrojadocharis Mattf.
  • Bahianthus R.M.King & H.Rob.
  • Barrosoa R.M.King & H.Rob.
  • Bejaranoa R.M.King & H.Rob.
  • Bishopiella R.M.King & H.Rob.
  • Campuloclinium DC.
  • Catolesia D.J.N.Hind
  • Conocliniopsis R.M.King & H.Rob.
  • Dasycondylus R.M.King & H.Rob.
  • Diacranthera R.M.King & H.Rob.
  • Gyptidium R.M.King & H.Rob.
  • Gyptis Cass.
  • Lapidia Roque & S.C.Ferreira
  • Lasiolaena R.M.King & H.Rob.
  • Litothamnus R.M.King & H.Rob.
  • Lourteigia R.M.King & H.Rob.
  • Macropodina R.M.King & H.Rob.
  • Morithamnus R.M.King, H.Rob. & G.M.Barroso
  • Neocuatrecasia R.M.King & H.Rob.
  • Prolobus R.M.King & H.Rob.
  • Semiria D.J.N.Hind
  • Stylotrichium Mattf.
  • Tamaulipa R.M.King & H.Rob.
  • Urolepis (DC.) R.M.King & H.Rob.
  • Vittetia R.M.King & H.Rob.
네오미란데아아족(Neomirandeinae R.M.King & H.Rob.)[3]
  • Neomirandea R.M.King & H.Rob.
등골나물아재비아족(Ageratinae Less.)[4]
  • 등골나물아재비속(Ageratum L.)
  • Acritopappus R.M.King & H.Rob.
  • Ascidiogyne Cuatrec.
  • Cavalcantia R.M.King & H.Rob.
  • Conoclinium DC.
  • Ellenbergia Cuatrec.
  • Ferreyrella S.F.Blake
  • Guevaria R.M.King & H.Rob.
  • Macvaughiella R.M.King & H.Rob.
  • Microspermum Lag.
  • Nesomia B.L.Turner
  • Paneroa E.E.Schill.
  • Phalacraea DC.
  • Phania DC.
  • Piqueriella R.M.King & H.Rob.
  • Radlkoferotoma Kuntze
  • Scherya R.M.King & H.Rob.
  • Teixeiranthus R.M.King & H.Rob.
등골나물아족(Eupatoriinae Dumort.)[5]
  • 등골나물속(Eupatorium L.)
  • Austroeupatorium R.M.King & H.Rob.
  • Criscianthus Grossi & J.N.Nakaj.
  • Eutrochium Raf.
  • Hatschbachiella R.M.King & H.Rob.
  • Stomatanthes R.M.King & H.Rob.
디시나피아아족(Disynaphiinae R.M.King & H.Rob.)[6]
  • Acanthostyles R.M.King & H.Rob.
  • Campovassouria R.M.King & H.Rob.
  • Disynaphia DC.
  • Grazielia R.M.King & H.Rob.
  • Raulinoreitzia R.M.King & H.Rob.
  • Symphyopappus Turcz.
리아트리스아족(Liatrinae R.M.King & H.Rob.)[7]
  • Carphephorus Cass.
  • Garberia A.Gray
  • Hartwrightia A.Gray
  • Liatris Gaertn. ex Schreb.
  • Litrisa Small
  • Trilisa (Cass.) Cass.
물머위아족(Adenostemmatinae B.L.Rob.)[8]
  • 물머위속(Adenostemma J.R.Forst. & G.Forst.)
  • Gymnocoronis DC.
  • Sciadocephala Mattf.
미카니아아족(Mikaniinae R.M.King & H.Rob.)[9]
  • Mikania Willd.
아야파나아족(Ayapaninae R.M.King & H.Rob.)[10]
  • Alomiella R.M.King & H.Rob.
  • Ayapana Spach
  • Ayapanopsis R.M.King & H.Rob.
  • Condylidium R.M.King & H.Rob.
  • Gongrostylus R.M.King & H.Rob.
  • Gymnocondylus R.M.King & H.Rob.
  • Heterocondylus R.M.King & H.Rob.
  • Isocarpha Less.
  • Lepidesmia Klatt
  • Monogereion G.M.Barroso & R.M.King
  • Parapiqueria R.M.King & H.Rob.
  • Polyanthina R.M.King & H.Rob.
  • Siapaea Pruski
알로미아아족(Alomiinae Less.)[11]
  • Ageratella A.Gray ex S.Watson
  • Alomia Kunth
  • Asanthus R.M.King & H.Rob.
  • Austrobrickellia R.M.King & H.Rob.
  • Barroetea A.Gray
  • Brickellia Elliott
  • Brickelliastrum R.M.King & H.Rob.
  • Carminatia Moc. ex DC.
  • Condylopodium R.M.King & H.Rob.
  • Crossothamnus R.M.King & H.Rob.
  • Dissothrix A.Gray
  • Dyscritogyne R.M.King & H.Rob.
  • Flyriella R.M.King & H.Rob.
  • Goyazianthus R.M.King & H.Rob.
  • Helogyne Nutt.
  • Kyrsteniopsis R.M.King & H.Rob.
  • Leptoclinium (Nutt.) A.Gray
  • Malperia S.Watson
  • Planaltoa Taub.
  • Pleurocoronis R.M.King & H.Rob.
  • Pseudobrickellia R.M.King & H.Rob.
  • Steviopsis R.M.King & H.Rob.
오악사카니아아족(Oaxacaniinae R.M.King & H.Rob.)[12]
  • Oaxacania B.L.Rob. & Greenm.
옥실로부스아족(Oxylobinae R.M.King & H.Rob.)[13]
  • 서양등골나물속(Ageratina Spach)
  • Jaliscoa S.Watson
  • Jaramilloa R.M.King & H.Rob.
  • Kaunia R.M.King & H.Rob.
  • Oxylobus (Moc. ex DC.) A.Gray
  • Spaniopappus B.L.Rob.
  • Standleyanthus R.M.King & H.Rob.
크리토니아아족(Critoniinae R.M.King & H.Rob.)[14]
  • Adenocritonia R.M.King & H.Rob.
  • Amboroa Cabrera
  • Antillia R.M.King & H.Rob.
  • Aristeguietia R.M.King & H.Rob.
  • Asplundianthus R.M.King & H.Rob.
  • Austrocritonia R.M.King & H.Rob.
  • Badilloa R.M.King & H.Rob.
  • Bishovia R.M.King & H.Rob.
  • Castanedia R.M.King & H.Rob.
  • Chacoa R.M.King & H.Rob.
  • Ciceronia Urb.
  • Corethamnium R.M.King & H.Rob.
  • Critonia P.Browne
  • Critoniadelphus R.M.King & H.Rob.
  • Critoniella R.M.King & H.Rob.
  • Cronquistianthus R.M.King & H.Rob.
  • Eupatoriastrum Greenm.
  • Eupatorina R.M.King & H.Rob.
  • Fleischmanniopsis R.M.King & H.Rob.
  • Grisebachianthus R.M.King & H.Rob.
  • Grosvenoria R.M.King & H.Rob.
  • Hughesia R.M.King & H.Rob.
  • Idiothamnus R.M.King & H.Rob.
  • Imeria R.M.King & H.Rob.
  • Koanophyllon Arruda
  • Lorentzianthus R.M.King & H.Rob.
  • Malmeanthus R.M.King & H.Rob.
  • Mexianthus B.L.Rob.
  • Neocabreria R.M.King & H.Rob.
  • Neohintonia R.M.King & H.Rob.
  • Nothobaccharis R.M.King & H.Rob.
  • Ophryosporus Meyen
  • Peteravenia R.M.King & H.Rob.
  • Santosia R.M.King & H.Rob.
  • Sphaereupatorium Kuntze
  • Steyermarkina R.M.King & H.Rob.
  • Tuberostylis Steetz
  • Uleophytum Hieron.
  • Urbananthus R.M.King & H.Rob.
  • Viereckia R.M.King & H.Rob.
트리코고니아아족(Trichogoniinae V.L.Rivera, S.C.Ferreira & Panero)[15]
  • Platypodanthera R.M.King & H.Rob.
  • Trichogonia (DC.) Gardner
  • Trichogoniopsis R.M.King & H.Rob.
트리코코로니스아족(Trichocoroninae R.M.King & H.Rob.)[16]
  • Sclerolepis Monnier
  • Shinnersia R.M.King & H.Rob.
  • Trichocoronis A.Gray
프락셀리스아족(Praxelinae R.M.King & H.Rob.)[17]
  • Chromolaena DC.
  • Eitenia R.M.King & H.Rob.
  • Eupatoriopsis Hieron.
  • Lomatozona Baker
  • Osmia Sch.Bip.
  • Osmiopsis R.M.King & H.Rob.
  • Praxeliopsis G.M.Barroso
  • Praxelis Cass.
플레이스크만니아아족(Fleischmanniinae R.M.King & H.Rob.)[18]
  • Fleischmannia Sch.Bip.
  • Sartorina R.M.King & H.Rob.
  • Zyzyura H.Rob. & Pruski
피퀘리아아족(Piqueriinae Benth. & Hook.f.)[19]
  • 스테비아속(Stevia Cav.)
  • Carphochaete A.Gray
  • Centenaria P.Gonzáles, A.Cano & H.Rob.
  • Cronquistia R.M.King
  • Iltisia S.F.Blake
  • Metastevia Grashoff
  • Piqueria Cav.
  • Piqueriopsis R.M.King
헤베클리니움아족(Hebecliniinae R.M.King & H.Rob.)[20]
  • Amolinia R.M.King & H.Rob.
  • Bartlettina R.M.King & H.Rob.
  • Decachaeta DC.
  • Erythradenia (B.L.Rob.) R.M.King & H.Rob.
  • Guayania R.M.King & H.Rob.
  • Hebeclinium DC.
  • Revealia R.M.King & H.Rob.
호프메이스테리아아족(Hofmeisteriinae R.M.King & H.Rob.)[21]
  • Hofmeisteria Walp.

계통 분류 편집

다음은 2002년 파네로와 펑크(Panero & Funk)의 연구에 기초하고[22] 2014년 갱신된 국화과의 계통 분류이다.[23][24][25]

국화아과
 
 

도로니쿰족

 
 

금방망이족

 
 
 
 

금잔화족

 
 
 

왜떡쑥족

 
 
 

길뚝개꽃족

 
 

참취족

 
 
 
 
 
 

금불초족

 
 
 

아트로이스마족

 
 
 

페데아족

 
 
 

헬레니움족

 
 
 

기생초족

 
 
 

폴림니아족

 
 

페리틸레족

 
 

네우롤라이나족

 
 

천수국족

 
 

바히아족

 
 

카이낙티스족

 
 

마디아족

 
 

밀레리아족

 
 

해바라기족

 
 

등골나물족

 
 
 
 
 
 
 
 

각주 편집

  1. ↑ Cassini, Alexandre Henri Gabriel de. Journal de Physique, de Chimie, d'Histoire Naturelle et des Arts 88: 202. 1819.
  2. ↑ King, Robert Merrill & Robinson, Harold Ernest. Phytologia 46(7): 446. 1980.
  3. ↑ King, Robert Merrill & Robinson, Harold Ernest. Phytologia 46(7): 448. 1980.
  4. ↑ Lessing, Christian Friedrich. Synopsis Generum Compositarum 155. 1832.
  5. ↑ Dumortier, Barthélemy Charles Joseph. Analyse des Familles des Plantes 31. 1829.
  6. ↑ King, Robert Merrill & Robinson, Harold Ernest. Phytologia 39(3): 133. 1978.
  7. ↑ King, Robert Merrill & Robinson, Harold Ernest. Phytologia 46(7): 447. 1980.
  8. ↑ Robinson, Benjamin Lincoln. Proceedings of the American Academy of Arts and Sciences 49: 435. 1913.
  9. ↑ King, Robert Merrill & Robinson, Harold Ernest. Phytologia 46(7): 448. 1980.
  10. ↑ King, Robert Merrill & Robinson, Harold Ernest. Phytologia 46(7): 446. 1980.
  11. ↑ Lessing, Christian Friedrich. Synopsis Generum Compositarum 154. 1832.
  12. ↑ King, Robert Merrill & Robinson, Harold Ernest. Phytologia 46(7): 449. 1980.
  13. ↑ King, Robert Merrill & Robinson, Harold Ernest. Phytologia 38(4): 323. 1978.
  14. ↑ King, Robert Merrill & Robinson, Harold Ernest. Phytologia 46(7): 447. 1980.
  15. ↑ Rivera, Vanessa Lopes, Ferreira, Silvana C. & Panero, José L.. Phytotaxa 260(3): 298. 2016. doi 10.11646/phytotaxa.260.3.10
  16. ↑ King, Robert Merrill & Robinson, Harold Ernest. Phytologia 46(7): 446. 1980.
  17. ↑ King, Robert Merrill & Robinson, Harold Ernest. Phytologia 46(7): 448. 1980.
  18. ↑ King, Robert Merrill & Robinson, Harold Ernest. Phytologia 46(7): 447. 1980.
  19. ↑ Bentham, George & Hooker, Joseph Dalton. Genera Plantarum 2: 165, 172. 1873.
  20. ↑ King, Robert Merrill & Robinson, Harold Ernest. Phytologia 46(7): 448. 1980.
  21. ↑ King, Robert Merrill & Robinson, Harold Ernest. Phytologia 46(7): 449. 1980.
  22. ↑ Panero, J.L.; Funk, V.A. (2002). “Toward a phylogenetic subfamilial classification for the Compositae (Asteraceae)”. 《Proc. Biol. Soc. Wash.》 115: 909–922. 
  23. ↑ Panéro, José J.; Freire, Susana E.; Ariza Espinar, Luis; Crozier, Bonnie S.; Barboza, Gloria E.; Cantero, Juan J. (2014). “Resolution of deep nodes yields an improved backbone phylogeny and a new basal lineage to study early evolution of Asteraceae”. 《Molecular Phylogenetics and Evolution》 80 (1): 43–53. doi:10.1016/j.ympev.2014.07.012. 2017년 1월 3일에 확인함. 
  24. ↑ Zhi-Xi Fu; Bo-Han Jiao; Bao Nie; Tiangang Gao (2016). “A comprehensive generic‐level phylogeny of the sunflower family: Implications for the systematics of Chinese Asteraceae”. 《Journal of Systematics and Evolution》 54 (4): 416–437. doi:10.1111/jse.12216. 2017년 1월 23일에 확인함. 
  25. ↑ Funk, Vicki A.; Fragman-Sapir, Ori (2009). 〈22. Gymnarrheneae (Gymnarrhenoideae)〉. V.A. Funk; A. Susanna; T. Stuessy; R. Bayer. 《Systematics, Evolution, and Biogeography of Compositae》 (PDF). Vienna: International Association for Plant Taxonomy. 327–332쪽. 2016년 12월 27일에 확인함. 
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