- ↑ Devi, R. & Boguskaya, N (2009). “Rhodeus atremius”. 《The IUCN Red List of Threatened Species》. 2016년 8월 14일에 확인함.
- ↑ Suzuki, N. and S. R. Jeon. (1988). Development of eggs, larvae and juveniles of Rhodeus ocellatus from Ansong-river, Korea (Pisces: Cyprinidae), with a note on minute tubercles on the skin surface. Kor. J. Limnol., 21(1): 1–15.
- ↑ Takuya Miyake, Jun Nakajima, Norio Onikura, Shigetoyo Ikemoto, Kei’ichiro Iguchi, Akira Komaru and Kouichi Kawamura. (2010). The genetic status of two subspecies of Rhodeus atremius,an endangered bitterling in Japan. Conservation Genetics 12(2):383-400.