참취족

참취족(--族, 학명: Astereae 아스테레아이[*])은 국화아과이다.[1] 약 170속에 2,800여 종을 포함하고 있다.

Infobox plantae.png
참취족
Aster amellus
Aster amellus
생물 분류생물 분류 읽는 법
계: 식물계
(미분류): 속씨식물군
(미분류): 진정쌍떡잎식물군
(미분류): 국화군
목: 국화목
과: 국화과
아과: 국화아과
족: 참취족
Astereae
Cass., 1819
모식속
참취속(Aster L.)

하위 분류편집

그랑게아아족(Grangeinae Benth. & Hook.f.)[2]
  • Akeassia J.-P.Lebrun & Stork
  • Ceruana Forssk.
  • Colobanthera Humbert
  • Cyathocline Cass.
  • Dacryotrichia Wild
  • Dichrocephala L'Hér. ex DC.
  • Egletes Cass.
  • Erodiophyllum F.Muell.
  • Eschenbachia Moench
  • Grangea Adans.
  • Grangeopsis Humbert
  • Grauanthus Fayed
  • Gyrodoma Wild
  • Heteromma Benth.
  • Mtonia Beentje
  • Nidorella Cass.
  • Pilbara Lander
  • Plagiocheilus Arn. ex DC.
  • Welwitschiella O.Hoffm.
데이지아족(Bellidinae Willk.)[3]
라게노포라아족(Lagenophorinae G.L.Nesom)[4]
  • Keysseria Lauterb.
  • Lagenocypsela Swenson & K.Bremer
  • Lagenophora Cass.
  • Myriactis Less.
  • Novaguinea D.J.N.Hind
  • Pappochroma Raf.
  • Piora J.Kost.
  • Pytinicarpa G.L.Nesom
  • Rhamphogyne S.Moore
  • Sheareria S.Moore
  • Solenogyne Cass.
  • Thespis DC.
마카이란테라아족(Machaerantherinae G.L.Nesom)[5]
  • Arida (R.L.Hartm.) D.R.Morgan & R.L.Hartm.
  • Benitoa D.D.Keck
  • Corethrogyne DC.
  • Dieteria Nutt.
  • Grindelia Willd.
  • Haplopappus Cass.
  • Hazardia Greene
  • Isocoma Nutt.
  • Lessingia Cham.
  • Leucosyris Greene
  • Machaeranthera Nees
  • Olivaea Sch.Bip. ex Benth.
  • Oonopsis (Nutt.) Greene
  • Pyrrocoma Hook.
  • Rayjacksonia R.L.Hartm. & M.A.Lane
  • Stephanodoria Greene
  • Xanthisma DC.
  • Xanthocephalum Willd.
  • Xylorhiza Nutt.
망초아족(Conyzinae Sch.Bip.)[6]
  • 개망초속(Erigeron L.)
  • Aphanostephus DC.
  • Apopyros G.L.Nesom
  • Darwiniothamnus Harling
  • Hysterionica Willd.
  • Leptostelma D.Don
  • Neja D.Don
미국쑥부쟁이아족(Symphyotrichinae G.L.Nesom)[7]
  • 미국쑥부쟁이속(Symphyotrichum Nees)
  • Almutaster Á.Löve & D.Löve
  • Ampelaster G.L.Nesom
  • Canadanthus G.L.Nesom
  • Psilactis A.Gray
미역취아족(Solidagininae O.Hoffm.)[8]
  • 미역취속(Solidago L.)
  • 미역취아재비속(Euthamia (Nutt.) Cass.)
  • Acamptopappus A.Gray
  • Amphiachyris (DC.) Nutt.
  • Amphipappus Torr. & A.Gray
  • Bigelowia DC.
  • Brintonia Greene
  • Chihuahuana Urbatsch & R.P.Roberts
  • Chrysoma Nutt.
  • Chrysothamnus Nutt.
  • Columbiadoria G.L.Nesom
  • Cuniculotinus Urbatsch, R.P.Roberts & Neubig
  • Eastwoodia Brandegee
  • Gundlachia A.Gray
  • Gutierrezia Lag.
  • Gymnosperma Less.
  • Lorandersonia Urbatsch, R.P.Roberts & Neubig
  • Medranoa Urbatsch & R.P.Roberts
  • Neonesomia Urbatsch & R.P.Roberts
  • Nestotus R.P.Roberts, Urbatsch & Neubig
  • Oligoneuron Small
  • Oreochrysum Rydb.
  • Petradoria Greene
  • Sericocarpus Nees
  • Stenotus Nutt.
  • Thurovia Rose
  • Xylovirgata Urbatsch & R.P.Roberts
바카리스아족(Baccharidinae Less.)[9]
  • Archibaccharis Heering
  • Baccharis L.
  • Commidendrum Burch. ex DC.
  • Melanodendron DC.
  • Microglossa DC.
  • Psiadia Jacq.
  • Psiadiella Humbert
볼토니아아족(Boltoniinae G.L.Nesom)[10]
  • Batopilasia G.L.Nesom & R.D.Noyes
  • Boltonia L'Hér.
  • Chloracantha G.L.Nesom, Y.B.Suh, D.R.Morgan, S.D.Sundb. & B.B.S
브라키스코메아족(Brachyscominae G.L.Nesom)[11]
  • Brachyscome Cass.
  • Calotis R.Br.
  • Ceratogyne Turcz.
아스트란티움아족(Astranthiinae G.L.Nesom)[12]
  • Astranthium Nutt.
  • Dichaetophora A.Gray
  • Geissolepis B.L.Rob.
  • Townsendia Hook.
참취아족(Asterinae (Cass.) Dumort.)[13]
  • 과꽃속(Callistephus Cass.)
  • 참취속(Aster L.)
  • Arctogeron DC.
  • Asterothamnus Novopokr.
  • Crinitaria Cass.
  • Galatella Cass.
  • Iranoaster Kaz.Osaloo, Farhani & Mozaff.
  • Kemulariella Tamamsch.
  • Miyamayomena Kitam.
  • Novopokrovskia Tzvelev
  • Psychrogeton Boiss.
  • Rhinactinidia Novopokr.
  • 갯개미취속(Tripolium) Nees
카이토파파아족(Chaetopappinae G.L.Nesom)[14]
  • Chaetopappa DC.
  • Monoptilon Torr. & A.Gray
크리솝시스아족(Chrysopsidinae G.L.Nesom)[15]
  • Bradburia Torr. & A.Gray
  • Chrysopsis (Nutt.) Elliott
  • Croptilon Raf.
  • Heterotheca Cass.
  • Noticastrum DC.
  • Osbertia Greene
  • Pityopsis Nutt.
  • Tomentaurum G.L.Nesom
펜타카이타아족(Pentachaetinae G.L.Nesom)[16]
  • Pentachaeta Nutt.
  • Rigiopappus A.Gray
  • Tracyina S.F.Blake
포도코마아족(Podocominae G.L.Nesom)[17]
  • Asteropsis Less.
  • Camptacra N.T.Burb.
  • Dichromochlamys Dunlop
  • Dimorphocoma F.Muell. & Tate
  • Elachanthus F.Muell.
  • Exostigma G.Sancho
  • Inulopsis O.Hoffm.
  • Iotasperma G.L.Nesom
  • Isoetopsis Turcz.
  • Ixiochlamys F.Muell. & Sond.
  • Kippistia F.Muell.
  • Laennecia Cass.
  • Microgyne Cass.
  • Microgynella Grau
  • Minuria DC.
  • Peripleura (N.T.Burb.) G.L.Nesom
  • Podocoma Cass.
  • Sommerfeltia Less.
  • Tetramolopium Nees
  • Vittadinia A.Rich.
호모크로마아족(Homochrominae Benth. & Hook.f.)[18]
  • Amellus L.
  • Chamaegeron Schrenk.
  • Chrysocoma L.
  • Engleria O.Hoffm.
  • Felicia Cass.
  • Gymnostephium Less.
  • Jeffreya Cabrera
  • Lachnophyllum Bunge
  • Nolletia Cass.
  • Poecilolepis Grau
  • Polyarrhena Cass.
  • Roodebergia B.Nord.
  • Zyrphelis Cass.
힌테루베라아족(Hinterhuberinae Cuatrec.)[19]
  • Achnophora F.Muell.
  • Aztecaster G.L.Nesom
  • Blakiella Cuatrec.
  • Cabreraea Bonif.
  • Celmisia Cass.
  • Chiliophyllum Phil.
  • Chiliotrichiopsis Cabrera
  • Chiliotrichum Cass.
  • Damnamenia Given
  • Diplostephium Kunth
  • Dysaster H.Rob. & V.A.Funk
  • Floscaldasia Cuatrec.
  • Flosmutisia Cuatrec.
  • Guynesomia Bonif. & G.Sancho
  • Haroldia Bonif.
  • Hinterhubera Sch.Bip.
  • Katinasia Bonif.
  • Laestadia Kunth ex Less.
  • Lepidophyllum Cass.
  • Llerasia Triana
  • Nardophyllum Hook. & Arn.
  • Novenia S.E.Freire
  • Ocyroe Phil.
  • Olearia Moench
  • Oritrophium (Kunth) Cuatrec.
  • Pachystegia Cheeseman
  • Pacifigeron G.L.Nesom
  • Parastrephia Nutt.
  • Pleurophyllum Hook.f.
  • Pteronia L.
  • Remya Hillebr. ex Benth. & Hook.f.
  • Rochonia DC.
  • Westoniella Cuatrec.
분류 불확실(incertae sedis)
  • Afroaster J.C.Manning & Goldblatt
  • Allittia P.S.Short
  • Apodocephala Baker
  • Aylacophora Cabrera
  • Doellingeria Nees
  • Ericameria Nutt.
  • Eucephalus Nutt.
  • Eurybia (Cass.) Cass.
  • Formania W.W.Sm. & J.Small
  • Hesperodoria Greene
  • Heteroplexis C.C.Chang
  • Heterothalamus Less.
  • Hullsia P.S.Short
  • Ionactis Greene
  • Madagaster G.L.Nesom
  • Mairia Nees
  • Nannoglottis Maxim.
  • Neobrachyactis Brouillet
  • Notopappus Klingenb.
  • Oclemena Greene
  • Oreostemma Greene
  • Pembertonia P.S.Short
  • Pseudobaccharis Cabrera
  • Rhynchospermum Reinw.
  • Roebuckiella P.S.Short
  • Sarcanthemum Cass.
  • Talamancaster Pruski
  • Toiyabea R.P.Roberts, Urbatsch & Neubig
  • Tonestus A.Nelson
  • Turczaninovia DC.
  • Vanclevea Greene
  • Vernoniopsis Dusén

계통 분류편집

다음은 2002년 파네로와 펑크(Panero & Funk)의 연구에 기초하고[20] 2014년 갱신된 국화과의 계통 분류이다.[21][22][23]

국화아과

도로니쿰족

금방망이족

금잔화족

왜떡쑥족

길뚝개꽃족

참취족

금불초족

아트로이스마족

페데아족

헬레니움족

기생초족

폴림니아족

페리틸레족

네우롤라이나족

천수국족

바히아족

카이낙티스족

마디아족

밀레리아족

해바라기족

등골나물족

각주편집

  1. Cassini, Alexandre Henri Gabriel de. Journal de Physique, de Chimie, d'Histoire Naturelle et des Arts 88: 195–196. 1819.
  2. Bentham, George & Hooker, Joseph Dalton. Genera Plantarum 2(1): 166, 176. 1873.
  3. Willkomm, Heinrich. Prodromus Florae Hispanicae 2: 30. 1870.
  4. Nesom, Guy Lane. Phytologia 76(3): 207. 1994.
  5. Nesom, Guy Lane. Phytologia 76(3): 208. 1994.
  6. Schultz, Carl Heinrich "Bipontinus". In: Webb, Philip Barker & Berthelot, Sabin. Histoire Naturelle des Îles Canaries 3(2.2): 217. 1844.
  7. Nesom, Guy Lane. Phytologia 76(3): 211. 1994.
  8. Hoffmann, Karl August Otto. Die Natürlichen Pflanzenfamilien 4(5): 144, 145. 1890.
  9. Lessing, Christian Friedrich. Linnaea 5: 145. 1830.
  10. Nesom, Guy Lane. Sida 19(2): 266. 2000.
  11. Nesom, Guy Lane. Phytologia 76(3): 203. 1994.
  12. Nesom, Guy Lane. Sida 19(2): 265. 2000.
  13. Dumortier, Barthélemy Charles Joseph. Florula belgica, opera majoris prodromus, auctore ... 66. 1827.
  14. Nesom, Guy Lane. Sida 19(2): 264. 2000.
  15. Nesom, Guy Lane. Phytologia 76(3): 203. 1994.
  16. Nesom, Guy Lane. Sida 19(2): 264–265. 2000.
  17. Nesom, Guy Lane. Phytologia 76(3): 209–210. 1994.
  18. Bentham, George & Hooker, Joseph Dalton. Genera Plantarum 2(1): 165, 174. 1873.
  19. Cuatrecasas, José. Webbia 24: 5. 1969.
  20. Panero, J.L.; Funk, V.A. (2002). “Toward a phylogenetic subfamilial classification for the Compositae (Asteraceae)”. 《Proc. Biol. Soc. Wash.》 115: 909–922. 
  21. Panéro, José J.; Freire, Susana E.; Ariza Espinar, Luis; Crozier, Bonnie S.; Barboza, Gloria E.; Cantero, Juan J. (2014). “Resolution of deep nodes yields an improved backbone phylogeny and a new basal lineage to study early evolution of Asteraceae”. 《Molecular Phylogenetics and Evolution》 80 (1): 43–53. doi:10.1016/j.ympev.2014.07.012. 2017년 1월 3일에 확인함. 
  22. Zhi-Xi Fu; Bo-Han Jiao; Bao Nie; Tiangang Gao (2016). “A comprehensive generic‐level phylogeny of the sunflower family: Implications for the systematics of Chinese Asteraceae”. 《Journal of Systematics and Evolution》 54 (4): 416–437. doi:10.1111/jse.12216. 2017년 1월 23일에 확인함. 
  23. Funk, Vicki A.; Fragman-Sapir, Ori (2009). 〈22. Gymnarrheneae (Gymnarrhenoideae)〉. V.A. Funk; A. Susanna; T. Stuessy; R. Bayer. 《Systematics, Evolution, and Biogeography of Compositae》 (PDF). Vienna: International Association for Plant Taxonomy. 327–332쪽. 2016년 12월 27일에 확인함.